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Leto's performance as a transgender woman in Dallas Buyers Club (2013) earned him an Academy Award, a Golden Globe Award, and a Screen Actors Guild Award for Best Supporting Actor.

Leto is considered to be a method actor, known for his constant devotion to and research of his roles.

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Their following releases, This Is War (2009), and Love, Lust, Faith and Dreams (2013), received further critical and commercial success. His parents divorced when he was a child, and he and his older brother, Shannon Leto, lived with their mother and their maternal grandparents, Ruby (Russell) and William Lee Metrejon.

Leto stated he "was raised around a lot of artists, musicians, photographers, painters and people that were in theater," adding that "Just having the art communal hippie experience as a child, there wasn't a clear line that was drawn.

Peter Stack of the San Francisco Chronicle noted how Leto played the athlete with raw vitality; "With hypnotic blue eyes and dirty blond hair, Leto captures the rock-star style Prefontaine affected, and he looks natural in fiery performances on the track, as well as off, where Pre affected a brash, confrontational style." After landing the lead role of a British aristocrat in the 1998 drama Basil, Leto starred in the horror Urban Legend.

The film was poorly received by most movie critics, however, it was a financial success.

To prepare for his role, Leto lived on the streets of New York City and refrained from having sex for two months prior to shooting.

His performance received critical acclaim by film critics who notably praised the actor's emotional courage in portraying the character's physical and mental degradation.

He made his film debut in How to Make an American Quilt (1995) and received critical praise for his performance in Prefontaine (1997).

Leto played supporting roles in The Thin Red Line (1998), Fight Club (1999) and American Psycho (2000), as well as the lead role in Urban Legend (1998), and earned critical acclaim after portraying heroin addict Harry Goldfarb in Requiem for a Dream (2000).

Leto returned to acting in 2002 with the thriller Panic Room, which marked his second teaming with director David Fincher.

He played the supporting role of Junior, a burglar who terrorizes Jodie Foster.

He often remains completely in character for the duration of the shooting schedules of his films, even to the point of adversely affecting his health.

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